Wolfgang Eberl wrote a special about this species group (DKG Journal 2016-1) I agree what he wrote: bochtleri and herzogi are confined to central Gabon (red oval). All others are related to herzogi. Some of them with very aberrant phenotypes and DNA. Complex group! - JVDZ
Type locality: Fast running brook about 3 kilometers north of Zoumoukou or 16 kilometers north of Lalara on the road to Mitzic and Oyem, northern Gabon, 0°27'N, 12°15'E.
